final Flashcards
temperate and boreal forests - geographic distribution
- predominantly in N hemisphere
- temperature forests do occur in S.America, Australia, and New Zealand
-in N hemisphere, coniferous forests form a broad circumpolar belt
temperate and boreal forests - climate
seasonal variation determined by temperature
TEMPERATE
- winter lows: -15 C to -5C
- frost free days; 120 to 150 days
- summer temps; 15 C to 27 C
- precipitation 500 to 1000 mm
CONIFEROUS
- winter lows: -30 C
- frost free days; 50 to 100 days
- summer temps; 12 C to 15 C
- precipitation <500 mm in summer and 100 mm snow in the winter
temperate and boreal forests - characteristic growth forms
TEMPERATE
- broad-leaved deciduous trees
- annuals and hemicryptophytes, forest floor
CONIFEROUS
- needle-bearing trees
- hemicryptophytes
temperate and boreal forests - characteristic species
TEMPERATE
- elm, ash, walnut, sugar maple, basswood, aspen, birch, maple, willows
CONIFEROUS
- white spruce, black spruce, jack pine, white pine, hemlock, cedar
comparison of the deciduous vs. the evergreen, coniferous habit – leaves
DECIDIOUS
- broad leaved
CONIFEROUS
- narrow, often needle-leafed
comparison of the deciduous vs. the evergreen, coniferous habit – tissue
DECIDIOUS
- vessels (perforate): wide diameter, efficient, competitive in mild climates due to efficient conduction
CONIFEROUS
- trachea’s (imperforate): narrow diameter, safe, competitive in seasonally dry environments due to ‘safety’ features
comparison of the deciduous vs. the evergreen, coniferous habit – photosynthetic rates
DECIDIOUS
- deciduous 15-35 mg CO2 dm-2h-1
-evergreen 15-18 mg CO2 dm-2h-1
-freezing results in cavitation
CONIFEROUS
- 5-18 mg CO2 dm-2h-1
-photosynthesis at temperature below freezing
temperate and boreal forests - adaptations to freezing low temps
plants cannot tolerate intracellular ice formation
- intracellular ice-crystal formation is avoided by:
(1) supercooling
(2) extracellular freezing
temperate and boreal forests - supercooling
-mestastable
- small cells
- low nucleation
- accumulation of solutes
- limit of -40 C
- wind –> ice
temperate and boreal forests - extracellular freezing
- plants decreasing in volume during freezing
- water of cell walls more dilute than cells
- as solvent extracted, depressed the freezing point
- cells potentially die of dehydration
- adaptations also at the membrane level (thawing expansion)
- lollipops, excision, fluidity
temperate and boreal forests - low temps and tree distribution
- trees species distribution is limited by low winter temperatures: both latitude and also elevation
- ring porous species restricted to milder regions (and new growth each year)
- diffuse porous species – not as limited, do not cavitate as easily in colder regions
DICOTS
a. constant size = diffuse porous
b. steady dec. in size = semi-ring porous
c. abrupt decrease in size = ring porous
CONIFERS
a. constant size
b. cell wall thickness increases gradually
c. cell wall thickness increases abruptly
temperate and boreal forests - treelines
at latitudinal extremes, conditions become too severe to support the growth of trees
- stress: physiological tissue damage due to low temperature or desiccation
- disturbance: mechanical damage due to wind abrasion, herbivory, snow loading, or fungi infection
- reproduction: reduced seedling and sapling establishment due to decreased pollination, seed development, seed dispersal, germination, or seedling establishment
- carbon balance: photosynthetic carbon gain minus respiratory demands is not enough to maintain minimum growth
- growth limitation: reduced development of new plant tissues due to low temps
the end of the tree-ed region may be abrupt, or represented by an ecotone of deformed trees: Krummholz
temperate and boreal forests- abrupt tree line
- shade tolerat deciduous species
- seedlings less tolerant of extremes than parents, establish under parents
temperate and boreal forests - krummholz tree line
- deformed tree scrub, includes coniferous species
- seedlings survive better than parents, but growth affected by environmental extremes resulting in Krummholz appearance
- Krummholz may be genetically fixed (?)
temperate and boreal forests - phenological responses
phenology in temperate and coniferous forests is in response to light : PHOTOPERIOD
- Coniferous forests slow down with decreasing light and temperature: reproduce in spring
- deciduous forests show obvious phonological responses to temperature and light
phytochrome
deciduous forests
- pigment in plants that mediates response to photoperiod
blue –> phototropins and cryptochromes
red –> phytochrome
- discovered through germination response of light requiring seeds
- exposure to red light –> germinate, far red light inhibits germination
- conversion of Pr to Pfr by red light, results in physiological responses such as seed germination
- ratios of Pr:Pfr and darkness are important in determining amount of active phytochrome and thus responses
R:FR at different times of day
daylight – 1.19
sunset – 0.96
moonlight – 0.94
deciduous trees - color change
color change in response to decreased photoperiod
- chlorophyll is broken down enzymatically
- carotenoids are unmasked
- anthocyanin are synthesized (produced as a result of altered sugar metabolism due to phosphate decrease)
- amino acids are stored till spring
deciduous tree leaf abscission
- reduce water loss, prevent damage from winter winds and snowfall, reduce insect predation
- hormonally due to IAA and ethylene
- development of an abscission zone seals off xylem, prevents further cavitation
leaf maintenance phase –> shedding induction phase –> shedding phases
deciduous tree reserves
accumulates starches (reserves for spring uses for new leaf flush) and sugars
deciduous tree bud break
occurs after a minimum cold period of one or more months followed by higher temperatures, 15 C to 20 C
- late flushing in oaks, etc., may ensure that transpirational demand remains low until new rings for conduction are formed
deciduous forests undestory
- therophytes germinate in spring
- stratification breaks dormancy of therophyes and cryptophytes
- changing light conditions on the forest floor lead to shade tolerant species becoming dominant in summer
temperate and boreal forests - sun and shade leaves
microclimates in the understories and in the tree canopies differ significantly in light
- there are shade tolerant and intolerant species
sun leaves (compared to shade) :
- thicker
- more stomata
-more soluble protein (Rubisco)
- more deeply lobed
leaf dimorphism also in conifers
temperate and boreal forests - fire
temperate deciduous forests seldom burn
- coniferous forests burn frequently
fire type: surface (needles) and/or crown
frequency: surface 1 - 10 years, crown 100 - 1000 years
intensity: low for surface fires, high for crown fires