Fish Flashcards
What are the general properties of fish?
- There are 20-30,000 species of fish
- Vast range in size, from less than a cm to 16m (whale shark)
- Found in wide range of habitats, including both fresh and salt water
- Mostly ectothermic (‘cold blooded’), can produce some heat via metabolism, but most of this is lost as the blood passes through the gills. This means that body temperature is usually within 1-2˚C of the water temperature, but there are exceptions.
- Efficient gill system to extract oxygen from water
- Fertilisation usually external, though some fish can bear live young
What is the Linnaean system of classification for rainbow trout?
Class: osteichthyes (have jaws, bone skeletons)
Sub-class: actinopterygii (membranous fins strengthened by bony rays)
Infra-class: teleostei (no ganoine in sclaes, upper jaw only attached to skull at snout)
Order: salmoniformes
Family: salmonid
Describe the outer anatomy of the eyes.
Eyes are large and do not have eyelids but the cornea is covered by transparent integument
Describe the outer anatomy of the ears and olfactory system.
No external ear is present in fish.
There are olfactory pits/nares which are used to detect scents in the water. There are 2 small openings rostral to the eye on both sides of the fish. Theses are connected by a small U shaped tube, which contains a vascular sensory epithelium. Water passes in through 1 opening and out the other and scents in the water are detected when water passes through the tube.
Describe the outer anatomy of the mouth and gills.
- Large mouth that is lined with teeth
- Maxilla is fixed cranially at the snout but free caudally. This is a feature of teleost fish.
- Operculum is a flap of skin covering the gills at each side of the caudal head. Supported by 4 thin bones. It is fixed cranially but free caudally and it can be lifted to expose the gills.
- Ventrally, the operculum is continued by a fold of skin called the branchiostegal membrane, which is supported by several small bones called the branchiostegal rays.
- Caudal edge of the operculum forms the boundary between the head and trunk.
List the features of the external anatomy.
Olfactory pit
Maxilla
Operculum
Lateral line
Pectoral fin - paired
Cranial dorsal fin
Pelvic fin - paired
Caudal dorsal/adipose fin
Anus/vent
Urogenital aperture
Anal fin
Caudal/tail fin
What is the position of the 2 openings on the trunk?
The trunk is marked by 2 openings on the ventral edge. Anus/vent is cranial while the urogenital aperture is more caudal.
What is the lateral line?
Lateral line is a dark line running along the length of the trunk. This continues as far as the caudal/tail fin. There are pits in this line containing sensory cells called neuromast cells. These have delicate hairs, protected by a gel-like cap and are used to detect movement and vibration in the surrounding water.
What is the collective purpose of the fins?
Fins are responsible for the fish’s ability to manoeuvre and remain stable in water. There are paired and unpaired fins.
Name and describe the 4 unpaired fins.
Cranial dorsal fin – on dorsal edge, protects the fish against rolling so stabilises the fish and assists in sudden stops and turns.
Caudal dorsal/adipose fin – soft and fleshy, absent in many fish but present in salmonids.
Anal fin – ventral edge, caudal to the urogenital aperture, used to stabilise the fish.
Caudal/tail fin – vertical, has species variations on shape, salmonids have homocercal tail where the fin is arranged symmetrically around the tip of the body. Tail is moved from side to side by the body muscles and propels the fish forwards.
Name and describe the 2 paired fins.
Pectoral fins – equivalent of forelimbs, for braking, balancing and turning, cranial.
Pelvic fins - equivalent of hindlimbs, for braking, balancing and turning, caudal.
List the layers of the integument of fish.
Cuticle/mucous/slime layer is outermost
Epidermis
Dermis
Scales
Describe the cuticle layer of the integument.
Has a protective function. Can make the fish more difficult to grab hold of so protects from predation, it reduces friction in water, contains anti-pathogenic immunoglobulins that act as an initial barrier to infection, slows down water transfer across the skin important for osmoregulation, retards decomposition of fish for food trade. If you catch a fish with the intention of returning it to water, you must be gentle so as not to disturb this layer. Excess mucous can be a sign of disease.
Describe the epidermis of the integument.
Cells retain capacity for cell division. Mucus secreting cells are abundant, producing the mucous to help form the cuticle layer. Relatively thin. Stratified squamous epithelium with mucous secreting cells between them.
Describe the dermis of the integument.
Divided into outer stratum spongiosum and inner stratum compactum. It contains chromatophores and scales, scales project towards the epidermis.
Describe the scales of the integument.
Calcified plates originating in the dermis and covered by the epidermis.
Name the 2 types of teleost scales.
Cycloid and ctenoid
Distinguish the 2 types of teleost scales.
Flat almost circular plates of bone that is laid down in concentric rings, which represent seasonal growth rates, so a fish could be theoretically aged with these
Cycloid scales have a smooth contour while ctenoid scales have stiff spikes protruding from one edge like a comb.
Describe the scales of the rainbow trout.
- Rainbow trout have cycloid scales
- Can have chromatophores associated with them.
- In trout, these are mainly melanophores that contain black or brown melanin granules, which can be concentrated or dispersed to allow colour to change.
- Dispersed granules give a darker colour as light disperses when passing through them.
What is cranial kinesis?
Maxilla is free caudally so movement is possible between the upper jaw and the cranium. This is cranial kinesis. Also occurs in reptiles and birds and is important to allow a wider gape.
Where are the teeth in teleost fish?
Most of the bones around the mouth cavity are equipped with teeth, which vary in shape and size depending on diet. Sharp pointed teeth indicate predatory feeding. Teeth can also be present inside the buccal cavity and lining the larynx and tongue.
Where are the branchial arches?
- Bones of the operculum are attached to the caudal aspect of the skull and overlap the bones of the branchial arches.
- Branchial arches are part of the gill apparatus and help form the wall of the pharynx.
- Each arch is a V shaped rod with the apex pointing caudally.
- There are 4 on each side of the fish.
- Dorsally and ventrally, the arches unite with others on the same side and those of the other side to form the pharynx.
Where is muscle distributed along the body?
- Complex musculature for head and fins
- Bulk of muscle is in trunk and tail. Muscle blocks arranged either side of the axial skeleton and attached to the vertebral column.
- Split into epaxial and hypaxial muscles
How is muscle arranged?
- Arranged in myomeres, zig-zag blocks. Each myomere has both cranial and caudal extensions, which extend under neighbouring myomeres. This overlap ensures the smooth transition of contractile force along the myomeres.
- Zigzag blocks separate by myosepta, which are connective tissue sheets.
- Coordinated contraction – undulations. Sequential contraction of muscle blocks allow movement.