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Paleobiology Flashcards

(744 cards)

1
Q

Paleontology / Paleobiology definition

A

study of ancient life

science of the forms of life existing in former geologic periods- represented by their fossils

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2
Q

Geology

A

geo- earth, logia/logos- study of

the study of Earth

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3
Q

studies earth material- minerals and rocks

processes operating within and on earth

A

physical geology

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4
Q

examines origin and solution of earths continents, atmosphere, oceans, and life

A

historical geology

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5
Q

system

A

combination of related parts that interact

biosphere, atmosphere, lithosphere, hydrosphere

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6
Q

determined that what are now mountains used to be the sea by the fact that mountain rocks contained shells

A

Leonardo da Vinci 1452-1519

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7
Q

demonstrated that fossils represent remains of ancient animals

A

Niels Stenson / Nicholas Steno (1638 - 1686)

shark teeth

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8
Q

previously called tongue stone

A

glossopetrae (sharks teeth)

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9
Q

recognized that fossil record showed species appearances and extinctions

A

Robert Hooke (1635-1703)

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10
Q

Georges Cuvier (1769-1832)

A

Father of vertebrate palaeontology and catastrophism

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11
Q

catastrophism

A

history of earth can be explained by sudden catastrophic events

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12
Q

uniformitarianism

A

the assumption that the same natural laws and processes that operate in the universe now have always operated in the universe in the past

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13
Q

fossil

A

the remains or impression of a prehistoric organism preserved in petrified form or as a mold or cast in rock

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14
Q

Why fossils are important

A
simple fascination
paleogeography
paleoecology
evolution
biostratigraphy
economics
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15
Q

compression

A

still contains parts of original organism

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16
Q

where can fossils be found

A

sediment, sedimentary rocks, metamorphic rocks, concretions/nodules

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17
Q

led to continental drift theories

A

paleogeography

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18
Q

paleogeography

A

environmental and physical restrictions to organism distribution

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19
Q

why fossils are important for evolution

A

fossils are the only direct record of the history of life

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20
Q

use of fossils in deducing succession and age relations, and dating sediments

A

biostratigraphy

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21
Q

index fossils must be

A
easily recognizable 
short stratigraphic range
easily preserved (hard parts)
worldwide/cosmopolitan distribution
rapidly evolving
abundant
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22
Q

an index fossil with a shorter stratigraphic range

A

gives more precise dating

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23
Q

economic fossils

A
chalk deposits
ammonites for building structure
dolomite mountains
fossil fuels ( coal, oil, gas)
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24
Q

rudist

A

cretaceous reefs with high porosity and permeability, important for reservoirs and caps

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25
process of fossilization | study of preservation
taphonomy
26
types of preservation
body fossils molds and cast ichnofossils
27
types of body fossils
unaltered remains | altered remains
28
unaltered remains happen
in unique environments where mechanical processes do not break down organisms
29
methods in which fossil organism may retain colour
freezing
30
types of unaltered remains preservation
Freezing Drying/ desiccation Amber/ tar/ wax/ asphalt
31
oldest fossil you could find from freezing
Quaternary
32
original materials for fossilization
CaCO3, Si, Apetite (calcium phosphate; teeth), chiton, cellulose, resistant organic substances (pollenin)
33
average ocean salinity
35ppt
34
photic zone
up to ~200m
35
freshwater salinity
<0.5ppt
36
brackish water salinity
5-30ppt
37
frozen fossils have
everything preserved - even internal organs
38
shore zone between high and low tide
littoral zone
39
below tide to edge of continental shelf, most diverse zone due to high nutrients and light
sublittoral zone
40
drifters, passive floaters
plankton
41
active swimmers
nekton
42
from continental shelf to abyssal plains
bathyal zone
43
benthic organisms on hard substrate
Epifaunal
44
benthic organisms on/in soft substrate
Infaunal (burrowing)
45
extract nutrients from sediment
deposit feeders
46
mobile marine organisms
vagile
47
immobile marine organisms
sessile
48
primary producers
cyanobacteria, algae
49
herbivores
gastropods
50
deposit feeders
gastropods, bivalves
51
suspension feeders
bivalves, crinoid
52
Amber fossils are mostly
insects from tertiary from baltic region
53
types of altered body fossils
permineralization recrystallization replacement carbonization
54
another name for permineralization
petrification
55
permineralization
porous material fills with groundwater original material not involved heavier than original material
56
most common permineralization
wood | Si fills pores
57
recrystallization
hard parts revert to more stable minerals or larger crystals
58
example of recrystallization
aragonite---- calcite
59
in dissolution and replacement, hard parts are replaced by
calcite, silica, pyrite, iron
60
pyritization occurs in
anoxic environments
61
example of dissolution and replacement
ammonite turns from iridescent/ mother or pearl -- pyrite
62
another name for carbonization
coalification
63
carbonization
only carbon remains, all other elements are removed, makes fossil appear black, organisms outline remains well preserved
64
common carbonization
organic rich shales, sandstones | plants (ferns), graptolites, fish
65
external / internal cavity
mould
66
filled in mould
cast
67
compression
2D shallow external molds that often display plant structures. retain original/chemically unaltered organic materials
68
impression
2D, no organic material, found in fine-grained sediment like clay or silt, commonly trace fossils, give insight into biological activity of organisms
69
tell more about organism behaviour than about the organism
trace fossils
70
types of trace fossils
``` tracks trails exogenic /endogenic trace fossil coprolite gastrolite ```
71
study of trace fossils
ichnology
72
vertebrate trace fossils
tracks
73
invertebrate trace fossils
trails
74
exogenic trace fossils
made on the surface of sediment (tracks)
75
endogenic trace fossils
made within the sediment (burrows)
76
fossilized poop
coprolite
77
fossilized digestive stones + stomach contents
gastrolites
78
only really distinct coprolite
sharks - swirly
79
coprolites & gastrolites tell
about diet not much else
80
set of trace fossils associated with a particular set of environmental conditions
ichnofacies
81
common / important ichnofacies
nereites, zoophycos, cruziana, skolithos, glossifungites, trypanites, teredolites
82
Nereites
deep marine ichnofacies | spiral, flower, sinuous, honeycomb
83
zoophycos
outer shelf - slope, low E muds, organic rich, | 3D feeding trace (burrow, tube)
84
cruziana
shallow marine shelf- upper slope | unidirectional, continuous, crawling trace
85
skolithos
``` shallow, near shore vertical burrows (dwellings) ```
86
Glossifungites
in firm - not lithified- sediment (mud, silt), marine intertidal and shallow plant root penetration, borings, burrows
87
Teredolites
borings in wood by bivalves
88
grazing traces are
very sinuous
89
resting fossil is
a depression in substrate
90
Trypanites
in hard substrate | predators (worms, bivalves, gastropods, barnacles) bore holes in corral, rock, shells
91
trace fossils characterized by behaviour
resting, dwelling, escape traces, moving, grazing, deposit feeding
92
pseudofossils
dendrites concretion / nodules solutioning
93
chemical / molecular fossils
chlorophyll, lipids molecule characteristic of cyanobacteria found before earth was oxygenated steranes evidence of eukaryotic life up to 1 billion years before they enter fossil records
94
dendrites
manganese, iron, tree like branching
95
how would bones be preserved
likely permineralization (porous)
96
how would teeth be preserved
likely recrystallization (solid)
97
preservation dependent on decay and mineralization
decay vs. mineralization, both must be minimum for soft parts to be conserved mineralized muscle--- tissue--- chitin--- cellulose--- shells
98
steps of taphonomy
necrolysis biostratinomy diagenesis
99
necrolysis
decay and decomp., from death to right before break up
100
biostratinomy
mechanical processes, before burial
101
diagenesis
mechanical, physical, chemical, and biological changes after burial
102
biocenosis
life assemblage, interacting organisms in a habitat
103
thanatocenosis
assemblage of organisms brought together after death
104
taphonomic filter
more and more information is lost with every step, some organisms are lost throughout filter
105
most resiliant bones
lower jaw, skull
106
physical diagenesis
compaction, deformation
107
chemical diagenesis
dissolution, recrystallization, replacement
108
biological diagenesis
bacterial decomposition (early stages)
109
how settling of bivalves can tell about environment
if they land concave up and stay that way it must have been a low energy environment
110
elongate minerals/fossils settle in what direction
parallel to shore
111
important for determining compaction/deformation
cleavage- to not mistake as elongate fossils
112
missing from fossil record
``` organisms without hard parts organisms from unpreserved environments organisms rare or geographically restricted habitat info behaviour living morphology ```
113
how organisms with hard parts can be lost from record
if soft parts are all that distinguishes them from organisms with similar hard parts
114
lost habitat info
ex. elephant bones found in a lake- they don't live there!
115
unpreserved environments
high latitudes (erosional environments)
116
communities lost from fossil record
only the biggest, strongest organisms are likely to survive, rare for young
117
material changed/moved due to erosion/reposition of sediment
reworking
118
reworking occurs by
lateral transport storm/currents bioturbation
119
examples of reworking
taxa ABC are eroded together- appear to have lived at same time but did not 2 halves of bivalve end up on different ends of beach kelp beds end up on beach sedimentation rate is so low that taxa ABC are deposited basically together
120
when a taxa disappears and then reappears although it never died
Lazarus taxa
121
lazarus taxa occurs due to
preservation issues environmental changes species endangerment
122
zombie effect
taxa 'reappears' after it has become extinct due to reworking
123
example of zombie effect
if an ammonite fossil surfaced now, but was reburied, and then resurfaced in the future
124
Elvis taxa
impersonating, morphologically similar organism to those before
125
Important for determining zombie effect
heat/colour alteration- sequence of colour change is irreversible, determines true age
126
how complete is the fossil record
>97% is not preserved
127
preservation favours which environment
marine (and some aquatic)
128
'motherload' of fossils
Lagerstratten
129
two types of Lagerstratten
Konzentrat | Konservat
130
Concentration deposits
Konzentrat concentrated but not best preservation large number of fossils, low preservation of minute details
131
types of Konservat
Stagnation deposits Obrution deposits Conservation traps
132
Konservat deposits
best type of preservation, fine minute details are preserved
133
Stagnation deposits
stagnant water supersaline lagoonal limits bacterial degredation ('pickled')
134
Conservation trap
Amber tar pits low decomposition
135
Obrution deposits
Organisms from normally not preserved environments are preserved Burgess Shale- large chunk of shelf slid deeper
136
Biasis of fossil record
``` rapid burial anoxic/hypersaline condition no/minimal reworking or diagenesis tissue resistant to decay organisms in low energy environment marine organisms ```
137
grouping of objects or information based on similarities
classification
138
taxonomy
branch of biology concerned with the grouping and naming of organisms based on their similarities, chemical make up, similarities
139
by observing patterns we can
deduce factors that control organism distribution
140
six major kingdoms represented by fossil record
``` Plantae Fungi Animalia Protista (uni/multi cellular) Archaeobacteria Eubacteria ```
141
extremophiles
Archaebacteria- live in harsh environments (anaerobic, hyper saline, sulfurous hot spring)
142
cyanobacteria
filamentous, less tolerant than Archaebacteria
143
Protists
protozoans- dinoflagellates,diatom, foraminifera
144
Fungi act as
decomposers | parasites
145
kingdom Fungi includes
molds mildew mushrooms yeast
146
plant evolution
green algae--spore plants (mosses)---vascular plants (ferns)--- seed plants (gymnosperms, angiosperms)
147
living organisms are classified by
binomial system of nomenclature | Linnaean system
148
Species
can interbreed and produce viable offspring basic unit of classification organisms with structural, functional, developmental similarities have unique two-part names
149
capitalized
genus name not species
150
italicized or underlined
genus and species
151
Classification hierarchy for plants
``` Kingdom Division Class Order Family Genus Species ```
152
classification hierarchy for animals
``` Kingdom Phylum Class Order Family Genus Species ```
153
mnemonic for classification hierarchy for animals
King Phillip Came Over From Great Spain
154
Paleontological species
based on similarities in morphology rather than genetic compatibility
155
Allopatric speciation
speciation that occurs when populations become isolated to an extent that prevents genetic interchange
156
sympatric speciation
one population slowly diverges from main population while still living in same area- don't interbreed
157
Which speciation is easier to deal with in fossil record
Allopatric
158
Allopatric speciation can occur either
symmetrically or asymmetrically
159
Phyletic gradualism
theory that evolution is gradual at a ~constant rate | speciation occurs as slow gradual change
160
punctuated equilibrium
theory that periods of evolutionary equilibrium are interrupted by episodes of rapid evolutionary change rapid change-- relative stasis-- rapid change--...
161
lineages show little evolution
punctuated equilibrium (stasis)
162
evolution takes place in lineages
phyletic gradualism
163
speciation is a side effect of evolution
phyletic gradualism
164
change in individual morphology during lifespan
ontogenesis | can compare species embryos
165
group of individuals of same species in an area
population
166
all populations of all species living in an area
community
167
a community and its abiotic environment
ecosystem
168
types of variation in organisms populations
ecophenotypic variation taphonomic variability sexual dimorphism copes rule
169
ecophenotypic variation
change due to environment- nutrient, light, temperature
170
taphonomic variability
distortion after death (biocenosis vs. thenatocenosis)
171
biocenosis
describes the interacting organisms living together in a habitat
172
thanatocenosis
death assemblage
173
sexual dimorphism
phenotypic variation between males and females of species
174
cope's rule
body size increases during evolution of a group of animals
175
types of skeletal growth
accretion addition molting modification
176
Accretion skeletal growth
adding on discrete growth layers to the skeleton as organism grows ex. corrals add layers every day, bivalves
177
recognizing sexual dimorphism
analogs to living organisms ex.antlers geographic time/location- F/M wouldn't be thousands of years apart in record Ratio ~50/50 M/F similarity at early life stages
178
Addition skeletal growth
adding discrete new parts which grow very little after formed ammonite, foraminifera (chambers)
179
Modification skeletal growth
continuous remodeling and adding to existing skeletal elements- mammals
180
Molting skeletal growth
shedding of exoskeleton- trilobite, crabs
181
Steno's three laws for sedimentary rocks
Principle of superposition Principle of original horizontality Principle of original lateral continuity
182
Physical principles of relative age
Principle of superposition, original horizontality, original continuity, cross cutting relations, inclusion, recorded history, unconformities, fossil succession, fossil correlation
183
principle of superposition
youngest strata is on top
184
principle of original horizontality
sediment is deposited in horizontal layers due to gravity
185
principle of original lateral continuity
continuity is preserved from one environment to the next
186
principle of recorded history
using known facts to date (ex. volcano eruption)
187
1986 peak in Cs level
chernobyl
188
principle of unconformities
surfaces of erosion or non-deposition (hiatus) include significant amounts of geologic time
189
principle of fossil succession
oldest fossils in a series of sedimentary rock layers will be found in the lowest layer
190
principle of fossil correlation
similar assemblages of fossils are of similar age and therefore the strata containing them are of similar ages
191
chronology of events in Earth history established on basis of obtaining ages of past events
Geological Time Scale
192
chronometers
radioactive elements | can measure up to ~7 half lives
193
if radioactive element has long half life
not able to measure daughter products in young materials
194
if radioactive elements have short half lives
cannot measure daughter product beyond certain time
195
elements for absolute dating
Pb210 | Cs137
196
Pb210
good for <120year old sediment | half life 22.3 years
197
Eons
Phanerozoic Proterozoic Archean Hadean
198
Phanerozoic
visible life
199
Proterozoic
early life
200
Archean
ancient life
201
Hadean
greek mythological hell (Hades)
202
~90% of earth history
Precambrian: Proterozoic + Archaen + Hadean
203
Era
precambrian paleozoic mesozoic cenozoic
204
Period mnemonic
camels often sit down carefully perhaps their joints creek TQ
205
geological time- periods
cambrian, ordovician, silurian, devonian, carboniferous, permian, triassic, jurassic, cretaceous, tertiary, quaternary
206
we no classify carboniferous as
upper- Pennsylvanian | lower- Mississippian
207
biozone
body of rock whose lower and upper boundaries are based on the range of one or more taxa
208
at well established geological time boundaries there are
golden spikes
209
what defines biozone boundaries
large extinctions
210
no fossil records or rocks of
Hadean
211
epoch time unit
early/middle/late
212
epoch time-rock unit
lower/middle/upper | Lower Devonian rocks represent Early Devonian time
213
history of Earth
4.6 billion years
214
time of Hadean
4.6-3.8bya
215
time of Archaen
3.8-2.5bya
216
time of Proterozoic
2.5bya
217
time of Cambrian
542mya
218
time of Silurian
444mya
219
time of carboniferous
360-250mya
220
time of mesozoic
250-65mya
221
time of tertiary
65-2.6mya
222
time of quaternary
2.6mya-present
223
time of cenozoic
65mya-present
224
by end of Archean
earth had an atmosphere, greenhouse gases, plate tectonics, life
225
evidence of Precambrian life
morphological fossils (black chert) stromatolites chemical fossils
226
chemical fossils
C12/C13 ratios, organisms preferentially take up lighter C12 | Pristane/phytane evidence of photosynthesis
227
prokaryote size
~10µm
228
how long was all life on earth bacterial
~2bya
229
beginnings of life on earth
Isua formation of greenland Warrawoona group fig tree formation gunflint cherts
230
Isua Formation of Greenland
3.85bya, oldest altered sedimentary rocks, geochemical indicators, carbon isotopes
231
Warrawoona Group
Apex chert, Western Australia, 3.5bya, stromatolites, six types of filaments
232
Fig Tree Formation
3.4bya, South Africa, cyanobacteria filaments associated with stromatolites, light carbon isotope ratios, pristine and phytane in cherts
233
Gunflint cherts
Canada, 2.1bya, stromatolites, black cherts, chemical fossils, diverse prokaryotes, numerous bacteria types
234
oncolite
similar to stromatolites, instead of forming columns, they form spherical structures, often form around central nucleus, shell fragment, and calcium carbonate structure is deposited by encrusting microbes
235
most likely origin of life
hydrothermal vents (chemoautotrophs)
236
first life
early archaen
237
why hydrothermal vents are a likely source of life
abundant energy/mineral supply ocean protects from UV many prokaryotes live near vents
238
needed for life to occur
cellular structure metabolic assimilation of energy reproduction heredity
239
origin of life theories
creation extra terrestrial spontaneous generation inorganic model
240
inorganic model
genetic material evolved first in association with clay minerals and organic compounds were involved only later
241
hydrogen oxidation
2H2 + O2 ---- 2H2O + energy
242
sulfur reduction
S + H2 -- H2S + energy
243
methane production
CO2 + 4H2 -- CH4 + 2H2O + energy
244
first photosynthesizer
cyanobacteria
245
kingdoms in Archean
archeobacteria | eubacteria
246
free oxygen (Tertiary atmosphere)
2.5-26mya
247
BIFs
interbedded chert and iron rich minerals (Fe sulphides, Fe carbonates) need low level oxygen (precambrian)
248
BIFs first appear
3.8BYA (more common in proterozoic)
249
BIFs rare after
1.9BYA
250
indications of higher levels of oxygen
stromatolites | eukaryotes
251
oxygen 'sinks' start to fill up and O2 can accumulate in atmosphere
~2BYA
252
Proterozoic eon life
2.5-0.5BYA evolution of complex eukaryotes first multicellular life evolution of sexual reproduction
253
origin of eukaryotes
2-1.8BYA
254
origin of sexual reproduction
1.1BYA
255
origin of multicellular life
~0.7BYA
256
evidence of Proterozoic evolution
gunflint chert | bitter springs chert
257
Rodinia
supercontinent 'motherland' | 1.2BYA-600MYA
258
Breakup of Rodinia
snowball/ slush ball earth | extinction- glaciation prevents light
259
Ediacara fauna
``` also Vendian (soft-bodied animals) developed movement, symmetry very thin, up to 1m some anoxic species- maybe symbiosis possibly leathery no predation, filtering/grazing ```
260
Ediacaran phytoplankton
Acritarchs- main primary producer after cyanobacteria organic walled, unknown affinity resistant to dissolution
261
Cloudina sp.
first skeletal fossil (Ediacaran) CaCo3, CaPO4, tube dwelling worm Pre-late Cambrian
262
Tommotian Fauna
``` small shelly fossils CaPO4 first evidence of predation (protective pieces) evidence of competition (grew taller) 1-5mm ```
263
Cambria diversification
``` continents split up, new ecological niches, sea level changes, expanded cont. shelf, warm water, transgression all phyla (except bryozoa) appear VERY rapidly ```
264
transgression
sea level rises- shoreline moves inland- mud deposited directly on old beach sand
265
cambrian life
many more shelled species decreased stromatolite abundance all phyla (except bryozoa) appear VERY rapidly most major invertebrate classes
266
hard part advantages
protection from UV- can move to shallow water prevents drying in intertidal pool predation protection support
267
hard part disadvantage
energy of molting
268
Archaeocyathid
sponges, appear in Tommotian (early Cambrian) extinct by mid Cambrian 1-3cm diameter, 15cm tall cone-in-cone structure, pores, central cavity calcareous skeleton benthic, sessile, colonial or individual
269
excellent indicator of early Cambrian
Archaeocyaths- only marine, benthic, sessile, passive filter feeders in 20-100m water depth of tropic carbonate shelves
270
Kingdom, phylum, subphylum of trilobite
Kingdom: Animalia Phylum: Arthropoda Subphylum: Trilobitomorpha Class: Trilobita
271
Trilobite time frame
``` only in Paleozoic extremely common in Cambrian and Ordovician early Cambrian- end of Permian 8/9 orders appeared in Cambrian 15,000 species ```
272
Trilobite exoskeleton
calcite
273
main index fossil of cambrian
trilobite
274
Major body parts of trilobite
Cephalon, Thorax, Pygidium | Pleural lobes, axial lobe
275
evolutionary trends of trilobites
greatly elongated to transverse (widened) thoracic segments increased (60) or reduced (2) reduction/loss of eyes
276
Effacement
adaptation related to a burrowing lifestyle | smoothing out of lines for burrowing (or streamlined swimming)
277
Spinosity
defensive/stabilizing adaptation | can tell modality from direction spines stick out
278
pelagic trilobite morphologies
extremely large eyes | streamlined bodies
279
olenimorph
forms associated with low O2, high sulfur benthic habitat | thin exoskeleton, increased thoracic segments, wide flat body, symbiotic behaviour
280
why wide flat body in olenimorph
larger surface area : water interface for material exchange
281
environment changes for trilobites mostly related to
temperature changes- largest effect in shallow water
282
Burgess Shale
mid Cambrian quick burial, anoxic conditions soft parts preserved ~93 soft bodied organisms
283
one of the first large predators (seen in Burgess shale)
Anomalocaris canadensis (sea horse kinda shape) can tell predator from distinct bite marks in trilobites up to 60cm
284
earliest chordate
Pikaia | ~5cm, nodochord, zigzag muscles attached to nodochord
285
Burgess shale organism that was incorrectly reconstructed (upside down)
Hallucigenia | ~1cm long, protective spikes on back
286
why explosion in Cambrian
``` few predators increased atmospheric oxygen sea level rise (many habitats) evolution of hard skeleton higher nutrient levels ```
287
parts of evolutionary faunas graph
mud grabbers--stationary filter feeders--mobile filter feeders--soft bodied organisms 1-5 large extinction plateau in cambrian- max # of species that can live on mud substrate, need grow taller evolution majority of species diversification at end of Cambrian
288
cambrian fauna
trilobite fauna | mainly trilobites, also echinoderms, brachiopods
289
paleozoic fauna
brachiopod fauna | brachiopods, bryozoans, graptolites, cephalopods, crinoids
290
Ordovician changes
``` 485-443mya north of tropics open ocean most land masses southern supercontinent, Gondwana O2 at modern levels flooding of continents ```
291
Ordovician radiation
150 families in Cambrian | 400 families in Ordovician
292
sediment changes from Cambrian-ordovician
Cambrian flat planar beds | Ordovician bioturbated, by upper Ordovician can't tell bedding planes
293
Tiering
increase height and depth in substrate
294
Conodonts
worm like forms with teeth, CaPO4, change colour in irreversible way, CAI, teeth not attached to jaw marine, free swimming, pelagic and nektobenthic predators, carnivores or scavengers
295
conodont phylum
EITHER Hemichordata or Chordata (uncertain)
296
conodont importance
extinct in Triassic, good for dating between triassic/jurassic, if find conodonts KNOW its not Jurassic
297
conodont morphotypes
coniform ramiform pectiniform
298
coniform
single cone or cusp with small basal cavity
299
ramiform
bars with central cusp and denticles
300
pectiniform
diverse forms bearing a platform and numerous denticles
301
graptolites name
graptos - written | lithos - rock
302
graptolite colonies
multiple filter feeders float/fixed rapid evolution index fossil for Ordovician
303
'arms' of graptolite
stipes
304
'spikes' of graptolite
thecae
305
length of both 'arms' of graptolite
rhabdosome
306
mid point of graptolite
sicula
307
graptolite phylum
Hemichordata
308
graptolite class
Graptolithina
309
Graptolite orders
Order Dendroidea | Order Graptoloidea
310
Dendroidea persistence
mid Cambrian - Carboniferous
311
Graptoloidea persistence
Ordovician - mid-Devonian
312
Graptolite features
``` exclusively marine rapid evolution abundant excellent biostratigraphic indices of Ordovician and Silurian Pelagic or benthic mostly preserved as thin carbon films ```
313
sponge phylum
Porifera | no symmetry, holdfast, can reassemble, pores, flagella, spicules
314
levels of complexity in sponges
Ascon-- Sycon-- Leucon | Ascon most common
315
sponge classes
Class Hexactinellida Class Calcarea Class Demospongia Class Sclerospongea
316
types of sponge spicules
Megascleres-monaxons, triaxons, tetraxons, polyaxon | Microscleres
317
Hexactinellida
Neoproterozoic- resent Always glass (siliceous) Six rays (at 90º in cubic symmetry) ~10cm, diverse, abundant, cool, deep water
318
Calcarea
``` Cambrian- resent calcareous or organic spicules- monaxon, tetraxon, star-shaped, tuning-fork shaped, all shapes significant late Paleozoic reef builders max. diversity in Cretaceous warm shallow water (above CCD) indicator of paleoclimate ```
319
Demospongia
Cambrian- resent 'everything else' siliceous spicules and/or protein spongin spicules- tetraxon, monaxon, knobby, desmas, irregular never 90º larger, more solid, heavier in deeper water, lighter in shallow water
320
Class Sclerospongea
``` Stromatoporoids spp. calcareous Ordovician-Cretaceous- or resent? thought to be extinct calcareous skeletons, domes >5m laminae look like stromatolite, less deformity canal with star shape on surface of bump ```
321
graptolite preservation
carbonization, pyritization fine grain sandstone/shale, low energy carbonization is flat- no internal structure pyritization can give 3D structure- fills up parts
322
evolution of graptolites
highest diversity in Ordovician | fewer branches down to 1 by silurian-devonian
323
types of sponge spicules
sponginin, silica, calcium carbonate
324
'other' reef builders
corrals
325
another name for phylum Cnidaria
Coelenterata
326
what Cnidaria means
cnidos- stinging nettle
327
Phylum Cnidaria classes
Hydrozoa- polyps, hydras Scyphozoa- jellyfish Anthozoa- sea anemones, corals, sea fans, sea pens
328
Class Anthozoa persistance
Precambrian-recent
329
Class Anthozoa orders
Tabulata Rugosa Scleractinia
330
Tabulata persistance
Ordovician-Permian | Prominent reef builders: S-D
331
Rugosa persistance
Ordovician-Permian | Reefs: S-D
332
Scleractinia persistance
mid Triassic- recent | prominent reef builders: late T-J-C
333
Order Tabulata
calcite structure, plates perpendicular to height, honeycomb appearance, continuous structure, ALWAYS colonial, small-absent septa
334
Order Rugosa
horn-shaped, solitary or colonial, mostly calcite, some aragonite, well developed septum, curl up from sea floor, accrete layer every day, 4 ray symmetry (6 in 4)
335
Rugosa + Tabulata
``` same time (o-p) same area no holdfast low energy unlikely to have had symbionts ```
336
Order Scleractinia
solitary and colonial, 6-ray symmetry (6 in 6), polyps fused into long, meandering rows, still present, only aragonite, some zooxanthellae
337
corral symbiont
zooxanthellae
338
corals as climate indicators
isotherm ~21º
339
Class Scyphozoa persistance
Precambrian- present
340
Sub-classes of Anthozoa
Octocorallians | Zoantharia
341
Phylum Cnidaria, Class Anthozoa, Sub-class Octocorallians, Orders
Scleractinia Rugosa Tabulata
342
calcite ocean Mg/Ca mole ratio
<2 High Mg Calcite 1-2 Low-Mg calcite 0-1
343
aragonite ocean Mg/Ca mole ratio
2-6
344
high spreading rates =
ridge volume large calcite seas Mg:Ca ~1
345
calcite sea persistence
Cambrian - mid Mississipian | Cretaceous-Ng
346
Aragonite sea persistence
Mid Mississippian- Cretaceous
347
Cambrian reef builder
Archaeocyathans
348
Ordovician to Permian reef builder
Tabulate corrals | Stromatoporoids
349
low spreading rate
small ridge volume | aragonite sea
350
lophophore
ciliated feeding structure near mouth
351
have a lophophore
bryozoans | brachiopods
352
Phylum Bryozoa persistence
Ordovician - present
353
Bryozoa characteristics
aquatic (marine/fresh), mostly interconnected colonies, small, sessile, lophophore, shallow water, aragonite or arag.-calcite mix or chitin
354
Phylum Bryozoa classes
Phylactolaemata Stenolaemata Gymnolaemata
355
Class Phylactolaemata persistence
Cretaceous- Recent
356
Class Stenolaemata persistence
Ordovician- recent
357
Class Gymnolaemata persistence
Ordovician- recent
358
phylactolaemata
freshwater, no skeleton
359
stenolaemata
marine, circular lophophore, tubular skeleton
360
gymnolaemata
circular lophophore, box-like/saclike skeleton
361
bryozoa shape dependence on environment energy
Discoid- high energy Branching- low (lagoonal) Encrusting- low-mod Fenestrated (spiral)- moderate
362
lamp shells
phylum brachiopoda
363
Phylum Brachiopoda persistence
Cambrian-present
364
Brachiopoda characteristics
two valves, solitary filter feeder, lophophore, marine, intertidal-abyssal (usually shelf), plane of symmetry bisects shell, sessile, bottom dweller, free-living or rooted (infernal/epifaunal)
365
brachiopod dorsal valve
brachial valve
366
brachiopod ventral valve
pedicle valve
367
smaller brachiopod valve
Brachial valve ('hole' for pedicle)
368
brachiopod muscles
adductor muscle | diductor muscle
369
opening/closing brachiopod valves
open when diductor muscle is tightened
370
Brachiopoda classes
Class Inarticulata | Class Articulata
371
Inarticulata
CaPO4 shells, no teeth along hinge, infernal, functional anus, common in Cambrian
372
how inarticulata move up and down in burrow
hydrostatic pressure
373
Articulata
Calcite shells, teeth in sockets along hinge, no anus, bilaterally symmetrical shells, dominate Ordovician
374
Paleozoic fauna
articulate brachiopods, stony and lacy bryozoans, graptolites, stromatoporoids, cephalopods, crinoids
375
stromatoporoids reef building
silurian-devonian
376
stromatolite reef building
Archean- proterozoic
377
First precambrian fossil ever discovered
Charnia_ lived deep in the ocean where light could not reach no mouth or gut but filtered nutrients/particles from water
378
Death assemblage
Thanatocenosis
379
Obrution
A rapid burial or smothering event
380
Life assemblage
Biocenosis
381
Parts of geological time scale from left to right
Eon Era Period Epoch
382
evolutionary faunas
cambrian fauna paleozoic fauna modern fauna
383
modern fauna
bivalve-gastropod fauna
384
modern fauna organisms
bivalves, gastropods, vertebrates, echinoids, crustaceans
385
mollusca
soft body
386
phylum mollusca presence
precambrian? Cambrian-recent
387
phylum mollusca morphology
usually elongate, bilateral symmetry, mantle secretes shell, radula, one way gut, muscular foot, concentrated sensory organs in head (except bivalve), eyes, usually gills, sexual reproduction
388
radula
minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters the oesophagus
389
mollusca mode of life
marine/freshwater/terrestrial
390
Phylum mollusca classes
``` Amphineura Scaphopoda Gastropoda Bivalvia Cephalopoda Rostroconchia ```
391
Class Amphineura
polyplacophora, monoplascophora | chitons
392
Amphineura characteristics
primitive, univalve shell, benthic, algal/bacterial grazers | Polyplacophora- 8 CaCO3 valves, girdle around valves
393
Amphineura strat range
Cambrian-Holocene
394
Class Scaphopoda
tusk shaped univalve shell open at both ends, lack gills, infernal, deposit feeder, rare in fossil record, few cm's
395
Scaphopoda strat range
Ordovician - recent
396
changes in class Scaphopoda
Carboniferous- long, skinny, dirty white Miocene- shorter, fatter, whiter, ribbed Eocene- shorter, black
397
Bivalve class
Pelecypoda (axe foot) | or Lamellibranchs
398
Bivalve features
two valves, calcite/aragonite, bilateral symmetry, indistinct head, distinct foot, tentacles and eyes absent, sexual reproduction, ligament
399
Bivalve strat range
Cambrian - recent
400
Bivalve environment
marine or lacustrine
401
Bivalve ligament
horny elastic structure joining two valves dorsally and acting as a spring that causes the valves to open when adductor muscles relax
402
umbo
most prominent, highest part of bivalve shell, usually contains the valve's beak, the oldest point (earliest formed) of the valve
403
determining left/right valve
labeled with respect to the anterior end of the bivalve, when the umbones are facing upward by viewing posterior (siphons) end
404
Bivalve classifications based on teeth and sockets
Taxodonts Heterodonts Isodonts Dysodonts
405
Taxodonts
numerous, small teeth, subparallel or radial arrangement
406
Dysodonts
small simple teeth, near edges of valve, teeth not distinct | Devonian- recent
407
Isodont
very large teeth, either side of central ligament pit
408
Heterodont
2-3 cardinal teeth below jumbo, elongated lateral teeth | most Tertiary-recent bivalves are of this type
409
Rudist
extinct bivalves 2 different sized/shaped shells sessile, solitary or reef like masses
410
Rudist bivalve strat range
Jurassic- Cretaceous
411
Bivalve life modes
mostly filter feeders Infaunal: Burrowing, boring Epifaunal: byssally attached, reclining, cemented, swimming
412
byssally attached
attached to substrate by byssal threads
413
reclining
lying immobile and unattached to sea floor
414
cemented
attached to substrate by secreted shell material
415
Burrowing bivalve morphology
equal size/shape valves 2 adductor scars equal in size distinct pallial line
416
Boring bivalve morphology
thicker, equal valves, cylindrical in X-section | some have ridges and stout spines, other tubular in form
417
semi infaunal bivalve
elongate, fan-like, reduced anterior area | ex. pen shells, modiolids, ark shells
418
Class Bivalvia, Subclass Pteromorphia, Genus Inoceramus
most diverse in Jurassic-Cretaceous | species ranged ~0.2-0.5Ma
419
number of gastropod species
40,000 - 100,000
420
gastropoda habitats
marine/lacustrine/terrestrial
421
Gastropoda features
mostly herbivores, some carnivores, univalved shell, distinctive head, eyes, tentacles, radula; 180º torted body (anus above head), sexual reproduction
422
Gastropoda strat range
Cambrian - present
423
gastropod means
stomache - foot
424
Gastropod shell geometry
trochospiral / planispiral
425
Trochospiral
shell coiled in more than one plane
426
planispiral
coiled in a single plane
427
Gastropoda columella
little column or pillar, central anatomical feature of coiled shell, often only visible when shell is broken, sliced in half, or x-rayed, runs from apex of shell to midpoint of undersurface
428
class Gastropoda subclasses
Prosobranchia Opisthobranchia Pulmonata
429
important bivalves for stratigraphy
Rudist | Inoceramus
430
likes like Rugosa, but no septa
Rudist bivalve
431
difference between Gastropod and Cephalopod
Cephalopods have partitioned shells (chambers)
432
meaning of Cephalopod
head-foot
433
Cephalopod characteristics
all predators, tentacles, eyes, partitioned shells, no or univalve shell, organized nervous system
434
cephalopod mode of life
marine, nektonic (jet propulsion)
435
Cephalopods of interest
nautiloids, ammonites, belemnites
436
cephalopod strat range
late cambrian- present | rapid initial evolutions
437
Phylum Molluska Class Cephalopoda subclasses
Nautiloidea Ammonoidea Coleoidea
438
Nautiloidea strat range
late Cambrian- recent
439
nautiloidea special features
simple suture pattern- straight to gentle curve, septate shell, well developed eye, leathery hood covers eye, siphuncle, jaws, radula, tentacles, hyponome, gills
440
first common large predator
Nautiloidea
441
septate shell
chambered
442
nautiloidea life mode
nektonic (0-600m)
443
protoconch
first chamber
444
conch
whole shell
445
phragmocone
external chambered shell | all chambers except living chamber
446
one chamber of conch
camera
447
separate camera in nautiloid conch
septum
448
septum
internal partition which separates the chambers
449
connection of septum to outer wall of conch
sutures (under top layer of shell)
450
tube connecting living chamber with all previous chambers of nautilus/ammonoid
siphuncle
451
Nautilus siphuncle
middle
452
siphuncle use
connects all chambers in order to change their buoyancy
453
living chamber
space between aperture and last septum
454
hyponome
expels water, locomotive force
455
nautilus 'shapes'
Orthoconic Cyrtoconic Gyroconic Nautilicone
456
orthoconic
totally straight, pointy shell
457
cyrtoconic
tip is more curved than orthoconic
458
gyrocone
curved shells, from single curve to multiple coils
459
Ammonoidea features
complex sutures, siphuncle, widespread, index fossil, chambered shell
460
ammonoid siphuncle
present at ventral side (not middle)
461
ammonoid strat range
Devonian - Cretaceous | **Important index fossil (pelagic)
462
if direction of aperture is up, a lobe in the suture would be a
U
463
if direction of aperture is down, a saddle in the suture pattern would be
n
464
Types of ammonoid suture patterns
ammonitic ceratitic goniatitic orthoceratitic
465
Ammonites have
ammonitic sutures
466
orthoceratitic
Cambrian- recent | practically straight suture lines
467
goniatitic
devonian - permian | smooth suture line, angular lobe/saddles
468
ceratitic
Carboniferous- Triassic | rounded saddles, serrated lobes
469
ammonitic
Permian - Cretaceous | complex lobes and saddles, dendritic
470
cephalopod shell morphology types (curving)
Involute Convolute Evolute
471
involute cephalopod shell
hiding the baby coils, living chamber covers initial chambers
472
convolute cephalopod shell
shell partly encloses preceding whorls | baby centre is partially exposed
473
evolute cephalopod shell
baby centre is completely exposed | living chamber does not cover any old chambers
474
shape of shell is important for
centre of gravity
475
ammonoid keel
thickening along outer margin | like a rim down the outer middle
476
ammonoid ribs/ornamentation
thickening of external shell in 'ribs' | not necessarily coinciding with sutures
477
ornamentation and keel are for
balance and protection
478
sexual dimorphism
microconchs | macrcoconchs
479
macroconch
female ammonoid, larger (for egg production)
480
microconch
male, smaller, long thin piece of shell at end of living chamber
481
aptychi
two-valved closing hatch (hood) on ammonite | calcite (ammonite shell is aragonite)
482
animals in Subclass Coleoidea
squids, cuddlefish, octopus belemnites
483
Coleoidea features
internal or lacking skeleton, orthoconic, marine, siphuncle, phragmocone, beak, gills
484
belemnites
2 parts: well preserved rostrum, phragmocone | bullet shaped, pointy, no chambers, siphon, typical middle groove
485
belemnite strat range
Devonian - Cretaceous | abundant Jurassic - Cretaceous
486
why belemnite fossils are usually found in large groups
large mortalities; anoxic conditions, inside of plesiosaurus
487
example of coleoidea
belemnite
488
the branch of palaeontology dealing with the study of microscopic fossils
micropaleontology
489
three main branches of paleontology
micropaleontology macropaleontology ichnology
490
what makes microfossils useful for stratigraphic correlation
abundance, abundance, abundance, widespread (large distribution), sediment cores are not wide (lower chance of macrofossil), preservable, short start range (high accuracy in dating), have distinct forms (identifiable)
491
2 main groups of microfossils
'plant like' | 'animal like'
492
plant like microfossils
diatoms, silicoflafellates, coccolithophores, dinoflagellates, acritarchs, pollen and spores
493
animal like microfossils
conodonts, ostracods, foraminifera, radiolarian
494
Kingdom Protista, autotroph divisions
Pyrrophyta Chrysophyta Cholorphyta
495
Division Chryophyta classes
Chrysophyceae Bacillariophyceae Coccolithophyceae
496
Bacillariophyceae
Diatoms
497
two groups of diatoms
Pennales | Centrales
498
diatom morphology
unicellular, planktonic, siliceous frustule, 2 valves, 20-200µm, autotrophic
499
diatom mode of life
every aquatic environment, benthic and planktonic (photic zone), individual or colonial
500
pennales diatom
bilaterally symmetric raphe: elongate fissure down middle mostly benthic Oligocene- Recent
501
centrales diatom
``` radially symmetric ornamentation: striations, pores, spines, punctuations mostly planktonic form chains Upper Jerassic?- Cretaceous- recent ```
502
diatom valves
epitheca | hypotheca
503
epitheca valve
larger, older, outer, girdle elements connected to it
504
hypotheca valve
younger, smaller, inner
505
diatom carbon fixing
20-25% of all planetary
506
class Chrysophyceae, order Silicoflagellineae
marine, flagellate, planktonic, photosyntehtic, opaline interior siliceous test, connected tubular structures
507
silicoflagellate start range
cretaceous - recent
508
silicoflagellate test shapes
ring like, hexagonal, hemispherical, lattice
509
division Chrysophyceae class coccolithophyceae
single celled, very small, 2-20µm calcitic plates (coccoliths), photosynthetic, nanofossils, semi-transparent
510
coccolithophore mode of life
planktonic, marine, photic zone, narrow temperature range- warm water, 45ºN-45ºS
511
coccolith function for coccolithophore
protection (bacteria, physical, predator) flotation/buoyancy (spherical shape) light regulation (reflect sunlight) biochemistry (may secrete calcite to expel byproduct, increase biochemical efficiency)
512
coccolithophore strat range
upper Triassic- Jurassic- recent | mass extinction at end of cretaceous, adaptive radiation in paleocene
513
major coccolith deposits
cretaceous chalk (calcareous ooze)
514
what does it tell you if you find only pelagic diatoms
greater depth (too deep for benthic photosynthesizers)
515
ooze
>30% biogenic Si/CaCO3
516
cause of equatorial band of siliceous ooze
radiolarians
517
can calcareous ooze be found in the abyssal?
yes at MOR, sea mount, etc.. if there are parts of ocean floor above CCD
518
affect of ocean acidification on coccolithophores
increases number of/thickness of coccoliths
519
CCD
carbonate compensation depth- below which calcareous skeletons dissolve as fast as they fall from above
520
Division Chlorophyta classes
Prasinophytes | Chlorophytes
521
Division Chlorophyta class Prasinophyta order
Acritarchs
522
Division Chlorophyta class Chlorophytes orders
Chlorococcales | Volvocales
523
micropaleontology
the branch of palaeontology dealing with the study of microscopic fossils
524
Autotroph divisons
Pyrrophyta Chrysophyta Chlorophyta
525
Acritarch size
average ~40µm | 20-150µm
526
Acritarchs are
organic-walled phytoplankton of unknown affinity | aquatic (marine)
527
Acritarchs are important for what time period
paleozoic (abundant on continental shelves)
528
the study of pollen, spores, and other similar structures (living or fossil)
Palynology
529
'other' similar structures in palynology
palynomorphs
530
palynomorphs
animal parts FOL algae
531
animal part palynomorphs
mandibles of polychaeta
532
FOL
foraminiferal organic linings
533
algae palynomorphs
freshwater green algae | dinoflagellates
534
middle cretaceous reef builder
there is no middle cretaceous
535
when writing the name of a geological time unit
don't forget to include eon, era, period, epoch
536
cross bedding represents
shallow marine environment
537
labelling a field map
location, formation, geological time | Muir Creek, Sooke Formation, upper Oligocene
538
y-shaped pollen scar
contact area- where they were connected
539
gymnosperm pollen
winged, looks like mouse head (conifer)
540
dinoflagellate composition
mainly organic, some calcareous
541
pyrrophyta
dinoflagellates - red tides
542
prior to diatoms, major primary producer
acritarchs, cyanobacteria
543
dinoflagellate characteristics
20-150µm auto/hetero/mixotrophic flagella - migration most abundant coastal phyto
544
dinoflagellates are most abundant when
summer, autumn- following diatom bloom
545
significance of dinoflagellates ability to migrate
the environment they are found in is the one they prefer to be in
546
hypnozygote
dinoflagellate cyst/ dinocyst resting cyst resulting from sexual fusion; commonly thick-walled, typically 15 to 100 µm in diameter, often resistant and made out of dinosporin
547
amroured dinflagellates
overlapping cellulose plates- armor- theca
548
visual difference between calcareous and organic walled dinoflagellates
organic- brownish | calcareous- whitish
549
organic walled dinoflagellate cyst strat range
triassic-recent
550
autotrophic dinoflagellates are mostly found
offshore (clear/transparent)
551
heterotrophic dinoflagellates are mostly found
near shore (darker/brownish)
552
proximity to shore =
proximity to nutrients
553
why sea level alters number/type of species
proximity to shore.. | higher sea level = larger shelf
554
why cenozoic species were not altered by sea level
limited by Si
555
Silica input is a function of
weathering/erosion
556
dinoflagellate strat range
different than cyst range Cambrian? - Recent flourish in Mesozoic
557
dinoflag paleoenvironmental reconstruction
``` coastal/oceanic proximity- sea level change- based on autotroph/heterotroph surface water T salinity (precipitations) paleoproductivity human impact (pollution, eutrophication) ```
558
paleozoic phytoplankton
acritarchs
559
mesozoic phytoplankton
coccoliths | dinoflagellates
560
cenozoic phytoplankton
coccoliths dinoflagellates diatoms
561
plant like microfossils
``` diatoms silicoflagellates coccolithophores dinoflagellates acritarchs pollen/spores ```
562
animal like microfossils
foraminifera radiolarians conodonts ostracods
563
Heterotroph subphylums
Sarcodina | Ciliophora
564
Sarcodina
protozoa that move and capture food by pseudopodia | Radiolarians, Foraminifera
565
Radiolarians strat range
mid Cambrian - recent
566
Radiolarian characteristics
only marine, heterotrophic, some symbiotic, 100-2000µm, siliceous, form chert
567
radiolaria 'parts'
apex, spine, pore, outer/inner sphere, chambered lattice shell, basal mouth
568
radiolaria orders
Spummelaria (circular) | Nassellaria (helmet shaped)
569
Spummelarian strat range
mid Cambrian - recent
570
Nassellarian strat range
Carboniferous- recent
571
deep water radiolarian strat
Silurian - recent
572
radiolarians compete for silica with
diatoms
573
radiolarian evolution
from shallow - deep | skeletons finer and less robust
574
Subclass Rhizopoda strat range
Foraminiferida | Cambrian - recent
575
benthic foraminifera
Cambrian - recent agglutinated or calcareous (not below CCD) some flatter, longer (less spherical)
576
planktonic foraminifera
Jurassic -recent ONLY calcareous bulbous- floatin spines- predation, surface area
577
Foraminifera characteristics
mostly marine, calcareous/agglutinated/organic
578
Foraminifera are characterized by
Test microstructure Test morphology Aperture type
579
Test microstructure
agglutinated calcareous tectin (not usually preserved)
580
agglutinated test
composed of grains or fragments of foreign material cemented by organism
581
calcareous test
hyaline (glassy/transparent) | porcelainous (white/opaque)
582
origin of biogenic silica
radiolarians, sponge spicules
583
foramin means
window (pores are like windows)
584
benthic foramin microhabitats graph
sediment depth vs. oligo/meso/eutrophic conditions deep infaunal taxa - don't live in oligotrophic, deep in meso, shallow in eutrophic (high rate of organic material falling) oligo- only epifaunal
585
oligotrophic
environments that offer little to sustain life
586
eutrophic
oversupply of nutrients, explosive growth of plants/algae- die, consume oxygen, hypoxia
587
foram test morphology
single chambered, uniserial, biserial, triserial, planispiral, trochospiral
588
uniserial
chambers added in a straight fashion (cone shaped with each successive chamber larger than the last)
589
biserial
chambers added in alternating fashion (conish/braid or horsetail)
590
triserial
chambers added every 120º spiral
591
planispiral
chambers coiled in single plane evolute or involute like typical ammonoid
592
trochospiral
chambers form cone (looks like a flower from the top)
593
earliest foraminifera were
benthic, agglutinated, single chambered
594
aperture types
open end of tube, radiate, loop shaped, terminal slit, umbilical, iteriomarginal, multiple, with phi aline lip, with bifid tooth, with umbilical teeth
595
large foraminifera
fusulinidae | nummulitidae
596
Fusulinids
rice grain shaped paleozoic shallow seas add chambers along long axis
597
nummulitids
coin shaped planispiral/involute tertiary
598
evolute/involute
space on both sides enclosed by last whorl is termed Umbilicus- wide umbilicus = Evolute, narrow umbilicus = Involute.
599
first appearance of foraminifera
early cambrian
600
first multi chambered foraminifera
late cambrian
601
mass extinction of forms
end of Permian | including fusilinids
602
first planktonic forams
early Jurassic
603
extinction of 1/2 of deep water benthic forams
end paleocene
604
modern benthic forams evolve
middle Miocene
605
foraminiferal inner lining
organic, chambers, some have this not all
606
benefits of studying foraminifera
small, abundant, geographically distributed, test changes, Camb-Recent, short reproductive cycle, trace elements preserved in test, subject to dissolution
607
benefit of short reproductive cycle
represent more precise conditions | more specific dating
608
benefits of being subject to dissolution
live at certain depths- more precise
609
foraminifera can be used to determine
geologic age environments pollution
610
determining environments
bathymetry
611
determining pollution
tests are altered with pollution/toxicity
612
first land plants
late Ordovician
613
basic alga parts
holdfast- anchor support - not needed (water) photosynthesizing parts - whole plant
614
basic land plant parts
roots- anchor, absorb water/minerals stem- support, photosynthesis cuticle/stomata- resist desiccation, allow gas exchange leaf- photosynthesis
615
difficulties moving to land
support, desiccation, reproduction, water, nutrients, temperature range, UV
616
plants evolved from
green algae- charophyceans
617
vascular tissues
transport water/nutrients | provide internal support
618
Bryophytes
nonvascular, lack roots, depend on moisture, unique habitat
619
bryophytes strat
on land- late Ordovician - Recent | existed in water in Camb
620
seedless vascular plants require
water for reproduction
621
seedless vascular plants
ferns, horsetails, club mosses | flagellated sperm need water
622
early seedless vascular plant
Psilotum- no leaves, no roots Cooksonia- spores, stomata, no leaves Rhynia- primitive root, sporangia
623
Ferns
need moisture for reproduction, roots, leaves, alternation of generations, spores
624
alternation of generations
multicellular haploid gametophyte, alternates with multicellular diploid sporophyte
625
largest Silurian plant
Baragwanathia
626
Baragwanathia
Lycopod, 30cm-1m, leaves, typical of silurian, spores
627
early-mid-late devonian changes in plants
mid- rise of lycopsids- spore bearing trees spores flammable- large increase of fires large increase of tree fall in swampy areas-- coal formation
628
first trees
Archaeopteris spore bearing, large Late Devonian, coal source
629
lycopsids
up to 30m, branches only at top, leaves similar to palm
630
calamites
Sphenopsids (horsetails) | jointed hollow stem, horizontal underground stem-bearing roots
631
gymnosperm
naked seed
632
Permian flora
gymnosperms- conifers, seed-bearing vascular plants
633
gymnosperm characteristics
2 cone types, fertilization independent of water, delayed sporophyte development, dormancy, seed dispersal, germination Permian - Recent
634
gymnosperm cones
Male- sperm- pollen grains | Female- embryonic seeds
635
examples of gymnosperms
conifers, ginkgo, cycads
636
meaning of angiosperm
covered seed
637
angiosperm strat
Early Cretaceous - Recent
638
angiosperm characteristics
flowering, fruits with seeds, dominant, grow-regenerate-reproduce faster, most successful/advanced plants, better at surviving grazing
639
angiosperms evolved from
ferns
640
insects coevolved with
angiosperms
641
evolution of leaves
microphylls megaphylls symmetry serration
642
microphylls
one vascular trace
643
megaphylls
branched vascular trace
644
increased leaf vein symmetry
equal strength, support, distribution of water/nutrients
645
gymnosperm present dominance
only in high latitudes
646
leaf serration
temperate climates- serrated margins | warm/humid climate- smooth margins
647
carboniferous- evolution of
gymnosperms
648
cretaceous - evolution of
angiosperms
649
oxygen spike in carboniferous
evolution of trees | aided in increase of fires
650
Lepidodendron
50m tall lycopsid, each part was named separately
651
largest oxygen spike since cambrian
late carboniferous
652
lowest CO2 level since Cambrian
carboniferous-permian boundary
653
Why do we see patterns of diversity in the fossil record
Total rock area (sampling) Paleontological interest Fossil preservation
654
Evolutionary faunas
``` Cambrian fauna (trilobite fauna) Paleozoic fauna (brachiopod) Modern fauna (bivalve-gastropod) ```
655
fraction of energy passed between trophic levels
~10%
656
similarities between evolutionary faunas
all 3 grew exponentially then plateaued in accordance with niche space
657
Example of how geology influences biology
formation of Pangea-- PT extinction
658
ocean chemistry - pangea
transition to aragonite sea | decrease in calcite dependent organisms
659
high spreading rates =
calcite seas
660
average species duration
0.5-10 million years
661
background extinction rate
~2-4.6 families per MYA
662
red-queen hypothesis
must keep running to stay in the same place- organisms must constantly adapt to avoid extinction
663
mass extinction is
geologically short intervals of intense species extinction at least 40% > 10 M families per year
664
mass extinctions we recognize
``` Late O Late D P-T boundary Late Tri K-T boundary ```
665
mass extinction causes
bolide, climate, volcanism, sea level, ocean chemistry, combinations etc
666
after mass extinction
empty niche spaces rapidly refilled- large increase in diversity
667
Late O extinction
one of coldest times- lots of ice, 2 pulses of extinction (cooling, warming), 60% of maine genera, 25% of families
668
cooling extinctions
sea level decrease, less niche space tropic species can't migrate to a warmer place shallow water species lose habitat
669
warming extinction
rising sea level, glacial melting, cold living species more greatly affected
670
isotopic fractionation
glaciers trap lighter O (16), ocean enriched in heavy O, ocean organisms enriched in heavy O glacial melting- light O returns, ratio ~constant
671
Late D extinction
cause unknown- cooling? meteorite?, mostly affected marine life- 22% marine families, collapsed massive reefs
672
P-T extinction
~245MYA, 95% marine species, 75% land species, most severe extinction, stromatolite increase, regression- loss of niches space
673
P-T ocean life
trilobites, tabulata/rugose corals extinct brachiopods, bryozoans, crinoids, ammonoids hit hard snails, clams, nautiloids less affected no reefs fro next 15MY first stromatolites in normal environment since O
674
P-T land changes
2 groups of therapsids survive 67% of amphibians extinctinct 30% insect orders extinct
675
P-T causes
plate tectonics, glaciation, ocean chemistry, volcanism, bolide, combination
676
P-T volcanism
siberian flood basalts-- SO2, CO2, block sunlight, cool earth-- CO2 buildup warms earth
677
P-T ocean chemistry
worldwide long term deep-sea anoxia, high pyrite (FeS2), from Pangea formation and current shutdown
678
P-T meteor impact
5-11km, wave of superheated vapour, dust in atmosphere, global cooling, glaciation
679
10 major biological advances to note
origin of life, eukaryotes/origin of sex, multicellularity, skeletons, predation, biological reefs, terrestrialization, trees/forests, flight, consciousness
680
Late Triassic extinction
breakup of pangea, low diversity, high oxidation, stromatolites, warm, condone extinction, large evaporite deposits
681
K-T extinction
``` increased fern spores- cooling? affected marine and terrestrial 60% plankton extinct ammonites, many marine reptiles extinct affected large animals more- dinosaurs, large marine reptiles ```
682
why larger animals didn't pass K-T extinction
higher requirements, lower abundances, slower reproduction
683
why K-T extinction
most likely bolide-- tsunami
684
K-T meteor
Chixulub - Yucutan peninsula of Mexico
685
evidence of K-T meteor
crater of same age, soot (fire), glass spherules (tektites), clay with Ir/Pt enrichment, tsunamites
686
Alvarez hypothesis (1990)
Found increased Ir level in sed. while identifying age of rock using nuclear weapon testing
687
What could Ir enrichment be
volcanic, meteor/rain of comets, cosmic rays from supernova
688
About 70% marine species became extinct
End O extinction
689
largest mass extinction 95% of marine
end P extinction
690
dinosaurs, marine reptiles, ammonites, belemnites extinct
end K extinction
691
most important land extinction- floral overturn >95%
end T extinction
692
extinction impacted trilobites, graptolites, echinoderms, brachiopods
end O extinction
693
extinction affected reef-dwellers, cerracitic ammonites, brachiopods, bivalves
end T extinction
694
extinction severely affected brachiopods, bivalves, foraminifera
end K extinction
695
extinction affected cephalopods, fish, corals
end D extinction
696
extinction marks boundary between dominance of paleozoic and modern fauna
P-T extinction (end P)
697
least understood mass extinction
end D
698
extinction most likely due to major meteorite impact in mexico
end K
699
extinction may relate to formation of Pangea
end P
700
extinction from sudden major glaciation
end O
701
extinction from volcanic activity that was a function of the breakup of pangea
end T
702
stratigraphy
study of the relations of stratified rocks, especially age relationships
703
correlation
determining age correspondence between rocks geographically seperated
704
methods of correlation
``` biostratigraphy lithostratigraphy magnetostratigraphy radiometric dating event stratigraphy sequence stratigraphy ```
705
sequence stratigraphy
global changes in sea level
706
The Principle of Fossil Correlation
similar assemblages of fossils are of similar age and therefore the strata containing them are of similar ages
707
Principles of biostratigraphic correlation
based on strat ranges of fossils rock units with same species must have been deposited during temporal range of that species biostrat units are divided into zones rock layers correlated to biostrat unit by presence of index fossils
708
index fossil
abundant- found easily morphologically distinct- distinguishable Geographically widespread- large-scale correlations wide range of environments- occur in many types of rocks narrow strat range- precise correlation preservable
709
fundamental unit of biostratigraphy
biozone
710
biozone
strat interval defined by occurrence of one or more fossil species/genera
711
biozones are defined by
International Stratigraphic Guide
712
local-range zone
teilzone
713
teilzone
extent of the unit in a certain place | between FAD and LAD of that taxon locally
714
taxon-range zone
between FAD and LAD of that taxon GLOBALLY
715
concurrent-range zone
intersection of the ranges of 2 or more taxa | ex. between FAD of one and LAD of another
716
interval zone
interval between 2 successive FADs OR | 2 successive LADs
717
assemblage zone
characterized by 3+ taxa
718
Oppel zone
special case of assemblage zone | defined by FAD/LAD of 1 taxon but characterized by additional taxa
719
Abundance zone
Peak/Acme/subset of teilzone | index species has high level of abundance locally
720
biostratigraphy is complicated by
Lazarus, Elvis, and Zombie
721
stratotype
type section- designated exposure of a named layered strat unit/boundary that serves as the standard of reference.
722
type locality
specific geographic locality where stratotype was described/named
723
type area
type region | geographic are that encompasses stratotype/type locality of strat unit/boundary
724
specific stratal sequence used for definition/characterization of stratigraphic unit being defined
stratotype
725
specific geographic locality where stratotype is situated
type locality
726
isochronous
events occur regularly, or at equal time intervals
727
diachronous
Varying in age from place to place
728
In regards to time, biozones should be
ISOchronous | NOT diachronous
729
why zones may not be isochronous
``` slow dispersal rate from time of origin barriers- ecological restrictions local extinction locally incomplete strat succession locally incomplete sampling local facies change preservation bias ```
730
ICS
international commission of stratigraphy
731
snowball/slushball earth time
Cryogenian period (850-635MYA)
732
snowball earth glaciations
Kaigas, Sturtian, Marinoan
733
end of snowball earth set the stage for
Metazoan life
734
why glaciation was triggered during snowball earth
supercontinent breakup- increased runoff- low CO2
735
snowball earth evidence
paleomagnetic data cap carbonate isotopic ratios BIF formation - low productivity
736
why it is now thought to have been slush ball earth
light had to have been able to penetrate to avoid reverting back to Archean biology
737
evolution of metazoan life dependent on
oxidation state of water | nutrients- glaciation = high erosion = nutrient flux (PO4)
738
main control on primary productivity
PO4
739
led to increased nutrients during snowball earth
glaciation = high erosion = nutrient flux (PO4)
740
altered oxidation state of marine waters during slush ball earth
nutrient flux-- glacial retreat-- algae radiation
741
Metazoans
all animals other than protozoans and sponges multicellular differentiated tissues (Tommotians/SSFs)
742
distinguishes metazoans from plants/algae
digest food in internal chamber | lack rigid cell wall
743
evidence of metazoan life
fecal pellets (early proterozoic, end of snowball period) amino acids, fractionation of C, isotopes (biomarkers) body fossils supplement evidence
744
why do body fossils only supplement the evidence of metazoan life
because they are not abundant enough to use as evidence alone-- first metazoans were mostly soft bodied