Stem Adaptations Flashcards

1
Q

What is the role of stems?

A

Support leaves, flowers and seed heads in positions that are most advantageous for the plants survival and its ability to compete with other plants and for its reproduction.

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2
Q

What do stems do?

A
  1. Put the leaves up into the light.
  2. Green stems can photosynthesize.
  3. They can adapt to protect or help plants to climb.
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3
Q

List stem adaptations

A
  1. Thorns (Crataegus crus-galli)
  2. Prickles (Rosa canina)
  3. Stinging hairs (Urtica dioica)
  4. Propagation (stolons: Fragaria x ananassa)
  5. Propagation (Rhizomes: Geranium endressii)
  6. Store nutrients (CORMS)
  7. Tendrils (Lathyrus odoratus)
  8. Petiole (Clematis)
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4
Q

Define an apical meristem

A

An apical meristem sits at the apex of a shoot and initiates cell growth, allowing the shoot to extend.

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5
Q

What is a dormant bud?

A

A dormant bud is at the side of a shoot,
typically in a leaf axil, and does not normally grow; however following loss of, or damage to, the apical meristem, it is capable of growing.

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6
Q

What is a twining stem?

A

stems which circle around enabling the stem to grasp a support, so enabling plants with this adaptation to climb up vertically.
Wisteria sinensis.

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7
Q

What is a stolon?

A

stems which stretch out horizontally and root when they touch the ground, so propagating the plant. Strawberry, Fragaria × ananassa

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8
Q

What is a corm

A

swollen underground lower parts of a stem, which enables a plant to store nutrients during a dormant season.

Crocus tommasinianus.

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9
Q

What is a thorn?

A

sharp outgrowths of a stem which enable a plant to attach its stems to bark or the stems of other plants and so climb; it can also be a defensive mechanism to prevent animal climbing.

Rosa canina.

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10
Q

Why is it often possible to propagate woody plants from stem cuttings but not herbaceous plants?

A

a. In order to propagate a plant, it must have cambium tissue. Vascular cambium is a secondary meristematic tissue that is only found in woody stems and provides the lateral growth of the stem. Herbaceous perennials do not have lateral growth and therefore, have no vascular cambium. Thus, herbaceous perennials are unable to be propagated with stem cuttings.
b. Fully grown herbaceous perennials do not have any cambium. However, when early in the year, the new shoots of a herbaceous perennial have a tiny amount of cambium in the base of the new shoot. If a cutting is taken at this point, it is possible to propagate.

MODEL:
Most woody plants have lateral cambium in their stems, which is a layer of tissue, where cells replicate and which allows the stem (twig or branch) to gradually thicken.

In many species, this cambium layer can be persuaded to grow roots and develop a new shoot, i.e. to form a new plant – which is what
we are doing when we take cuttings.

In herbaceous plants, the mature stems
does not have any lateral cambium, as it is essentially a flowering stem, which
is growing upwards, and not thickening. Without cambium, there is no hope
of new growth being initiated, and so cuttings are impossible.

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11
Q

Suckers on woody plants are sometimes regarded as a problem, but in other circumstances, they might be a boon. Why the difference of opinion?

A

a. A sucker is a shoot originating underground from the root or stem of a plant.
b. As a plant sends out multiple shoots and forms numerous new clones it may become an invasive and difficult plant to eradicate from the garden.
c. However, if you are planning a habitat restoration site, it may provide a good dense thicket which would be very advantageous for birds and wildlife.
d. Knowing the site and the future needs of the area will determine whether you wish to plant something that produces numerous suckers. An example of a plant with suckers would be Rhus typhina or Musa paradisaica.

MODEL:
Suckers are analogous to the rhizomes of herbaceous perennials. For the purposes of propagation they are very useful, but in most garden situation a mass of new growth around the base of a tree or shrub is seen as untidy, or
needs controlling, to limit the spread of the plant.

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12
Q

The most common and easiest way of propagating herbaceous perennials is by division.
What is division?
Describe using appropriate terms.

A

a. Division is the simplest method of propagation where parts of the plant that show totipotency are separated

b. In a bulb, such as an Allium sativa, cutting the bulb in half, with roots, leaves and stems exactly the same will produce a second plant. For plants that have multiple base stems, such as herbaceous perennials, division must include both the roots and the stems. An example would be Rudbeckia fulgida var. deamii
You should mention rhizomes, (eg. Rudbeckia) as the key organ enabling divitson of herbaceous plants , rhizomes show strong totipotency. In other cases root tissue is totipotent allowing division.

MODEL:
Division involves digging up a plant and dividing the underground plant tissue. In some cases this will be root tissue, but in most cases it will be rhizomes, i.e. stems, which we are dividing. Since rhizomes naturally produce new (adventitious) roots and shoots regularly, this is a very easy way to propagate these plants.

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13
Q

What might be the advantages, for environmental improvement, and for biodiversity, in having climbers growing up several storeys on buildings? What kind
of support systems might be needed to enable
climbing plants to grow up tall buildings?

A

a. Advantages during winter will help to insulate the house from the cold, winds and temperature fluctuations. In the summer, the climbers will protect the house from the extreme temperature of the sun, again providing insulation for the house.
b. IF the climber is deciduous the house can receive sunlight for warmth during the winter and in the summer when the plant is in full growth, provide shelter from the extreme heat.
c. Climbers can also create habitat for birds and pollinators.
d. An issue with climbers on the walls is that some plants must be checked often to make sure they are not growing under the eaves into the roof.
e. Support systems will depend on the climber used but mostly a trellis or vertical wires will do:
e.i. Hedera helix has adventitious roots that will cling to the wall so very little support is needed.
e.ii. Plants with tendrils, which are evolved leaves, will need some sort of trellis attached to the wall in order for the tendrils to cling onto the plant. An example of plants would be Vitis vinifera. Best to use a fan trellis to begin the growth up the wall.
e.iii. Plants with prickles will need the support of wires or a trellis for the plant to grow. The Rosa spp. Has prickles that will help to attach to the trellis, but the plant will need full support in order to climb.
e.iv. Plants such as Wisteria sinensis is a twining plant that twists around poles, gutters or anything to gain height, so once it has secured itself to a structure it will climb.

MODEL:
Climbing plants on buildings provide many of the advantages of trees: shade, evaporative cooling, habitat. They can help mitigate some of the negative impacts of the ‘urban heat island effect’, which will increasingly be a problem with climate change. Growing them on, or adjacent to, buildings can help
moderate temperatures inside the building, provide insulation and reduce solar radiation. They can also be visually exciting. However their weight can be considerable, especially when laden with snow or being buffeted by wind, so they need good supports. For anything higher than two storeys a structural
engineer should be involved. Twining climbers only need vertical cables, but many others need a grid like structure to grasp to enable them to climb.

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14
Q

The use of herbaceous perennials in gardens and in public spaces is increasing. Herbaceous plantings can take many different forms. An important functional use is as ground cover. What might be the most useful aspects of perennial species for effective ground cover? What would you look for when selecting plant species?

A

Ground cover herbaceous perennials are most effective when they spread rapidly and form a very consistent carpet, without gaps. Those under 40cms are particularly useful, anything taller than this can be blown about by wind
and begin to look untidy. Only a minority of perennials are suitable, notably those with rapidly spreading and frequently branching rhizomes, such as Bergenia spp. or Geranium macrorrhizum. These are also easy and cheap to propagate in the quantities needed.

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15
Q

Where is new tissue grown?

A

Apical meristem

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16
Q

Does a Herbaceous Perennial have lateral meristem?

A

NO, only woody has lateral meristem for secondary growth.

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17
Q

What is the core of a tree made of?

A

Dead xylem. As the tree grows laterally the xylem and phloem spread out and middle dies. Rings of the tree are dead xylem.

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18
Q

What must a plant need to vegetatively propagate?

A

CAMBIUM.
Cells that allow for growth is the site for potential rooting.

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19
Q

How to propagate a herbaceous perennial

A
  1. Division. Dig up and separate entire plant so root/leaves are together.
  2. Heel cuttings are often effective but TIME SENSITIVE. Must be base of new fresh shoot that has cambium.
  3. Adventitious roots form, therefore stem may become a new plant. Using a cutting of such a plant may well produce a clone.
20
Q

List climbing stem adaptations

A
  1. Petiole: Twists around and grabs hold Clematis ‘Princess Diana’
  2. Thorns Bougainvillea glabra
  3. Prickles Rosa canina
  4. Tendrils - in case of grapes. Vitis labrusca
  5. Adventitious roots Ilex aquifolium?????
  6. Rhizomes: Geranium endressii
  7. Suckers: Musa paradisaica
  8. Corm: swollen stem to hold nutrients Crocus sativa
  9. Tubers Solanum tuberosum
  10. Stolons Fragaria x ananassa or Ranunculus repens
21
Q

What’s the difference between a rhizome and sucker?

A

Suckers are young shoots that arise from the base of a plant, usually from the roots or the stem, whereas rhizomes are underground horizontal stems that produce roots and shoots.

22
Q

What is a stolon?

A

Overground reproductive stem that arches out. When it hits the ground they take root. Same as a runner. Fragaria x ananassa.

23
Q

Define Totipotency

A

cellular totipotency is the ability of a single plant cell to produce a whole new plant autonomously as a single process.

24
Q

Define etiolated

A

a process in flowering plants grown in partial or complete absence of light. It is characterized by long, weak stems; smaller leaves due to longer internodes; and a pale yellow color (chlorosis).

25
Q

What would be an effective groundcover that spreads?

A

Vinca spp. Periwinkle. Spreads via rhizomes so may become invasive.

26
Q

What would be an effective moisture loving plant for woodland areas?

A

Matteucia struthiopteris. FERN, spreads via rhizome.

27
Q

Define perennation?

A

Storage organ of a plant that stores starches, nutrients through the dormant season.

28
Q

What is the lateral meristem responsible for?

A

Secondary growth that replaces the epidermis in roots and stems. Secondary growth produces the girth of the plant.

29
Q

What kind of plants have minimal stems, and why?

A

Alpine and desert plants. Protection from extreme temperatures.

30
Q

What is a soft wood cutting?

A

A cutting that is taken from young material in active growth. It often roots quickly. The stem has few reserves so it doesn’t work well for plants that develop more slowly.

31
Q

What is a semi-ripe cutting?

A

This is a more mature cutting with growth beginning to harden. This is the most successful, taking several weeks to root. The stem has enough vigor and auxins to allow rooting.

32
Q

What is a hardwood cutting?

A

1 year old stem made in early winter just after leaf fall. Can take up to 1 year to root. Salixx is actually faster than most 1 year cuttings.

33
Q

What is layering?

A

Shoots that are held down to the soil to root. Cornus spp.

34
Q

What is air-layering?

A

Shoots wounded and surrounded by damp moss enclosed in plastic bag. Some Magnolia spp can do air layering.

35
Q

What is grafting?

A

For plants that don’t root from cuttings, shoots are joined to the root stem of a close relative. The two layers of cambium meld and grow together as one. It combines the root system of one and top growth of another.

Most fruit trees are grafted.

36
Q
A
  1. Pith
  2. Phloem
  3. Xylem
  4. Primary Phloem fibers
  5. Secondary phloem
  6. Primary phloem fibers
  7. Cork
    8.Cork cambium
    9.Phelloderm
  8. Periderm
  9. Bark
  10. Outer bark
  11. Inner bark
  12. Phloem ray
  13. Cortex
  14. Secondary xylem
  15. Primary xylem
  16. Vascular cambium
37
Q

Define periderm

A

Structure that consists of a cork cambium (Phellogen) with cork tissue (phellem) to the outside, and in some cases a layer of cells derived from and to the inside of the cork cambium called phelloderm.

38
Q

What is the function of the phelloderm

A

Functions to limit dehydration and block pathogens after the epidermis is disrupted by the onset of secondary growth.

39
Q

Define cork

A

Tissue dead at maturity generated from a cork cambium

40
Q

What is cork cambium

A

Layer that functions to produce cork, and in some cases, phelloderm. It is derived from the cortex in stems. Over time one cork cambium will be supplanted by another generated from parenchyma cells

41
Q

What is phelloderm

A

A layer of secondary tissue generated by the cork cambium to the inside.

42
Q

What is a primary tissue

A

Tissues generated from the growth of an apical meristem

43
Q

What is cambium

A

A lateral meristem. These growth cells increase the girth of the plant organ.

44
Q

What are secondary tissues?

A

Tissues generated from the growth of the cambium.

45
Q

Explain secondary growth

A

The growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to produce secondary phloem and xylem. The stem will thicken, while primary growth is growth that occurs as a result of cell division at the tips of stems and roots causing them to elongate.

Monocots usually lack secondary growth